Caves have long held a profound mystic in the psyche of humans throughout the ages. Whether functioning as the spiritual ?doorway to the underworld,? a place of habitation, a philosophical allegory, or a recreational thrill, caves have played a dynamic role in our lives and in the lives of countless other organisms. This course will begin by reviewing human interaction with caves over time and will transition to our modern understanding of the geology behind cave formation. We will then extend our scope beyond humans and work to understand how and why certain organisms call caves ?home.? Our analysis will highlight evolutionary processes that have lead to exhilarating and often peculiar adaptations found in organism that live in or near caves. We will also address current topics in cave biology including their use for studies of global climate changes and pervasive problems associated with human interference in these delicate ecosystems. This course will feature multiple field tr